HR job title
- Retired Professor
- Contact Info
- 1 979 845 3541
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Varner, Dickson Professor
Positions
Dr. Dickson Varner is a Diplomate of the American College of Theriogenologists. His special interests include equine theriogenology, fertility probes for stallions, in-vitro preservation of equine spermatozoa, capacitation of equine spermatozoa, assisted reproductive techniques, and subfertility in stallions.
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Overview
Publications
selected publications
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academic article
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Serafini, R., Ghosh, S., Love, C. C., Medrano, J., Teague, S. R., LaCaze, K. A., & Varner, D. D.
(2019).
Effect of artificial vagina lubricants on stallion sperm quality. THERIOGENOLOGY.
139, 121-125.
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Hernndez-Avils, C., Love, C. C., Serafini, R., Teague, S. R., & Varner, D. D.
(2019).
Supplemental Antibiotics in a Commercial Extender for Stallion Semen. 80, 33-35.
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Valdez, R., Cavinder, C. A., Varner, D. D., Welsh, T. H., Vogelsang, M. M., & Ing, N. H.
(2019).
Dexamethasone downregulates expression of several genes encoding orphan nuclear receptors that are important to steroidogenesis in stallion testes. Journal of Biochemical and Molecular Toxicology.
33(6), e22309.
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Hernndez-Avils, C., Serafini, R., Love, C. C., Teague, S. R., LaCaze, K. A., Lawhon, S. D., ... Varner, D. D.
(2018).
The effects of antibiotic type and extender storage method on sperm quality and antibacterial effectiveness in fresh and cooled-stored stallion semen. 122, 23-29.
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Serafini, R., Varner, D. D., Blanchard, T. L., Teague, S. R., LaCaze, K., & Love, C. C.
(2018).
Effects of seminal plasma and flash-freezing on DNA structure of stallion epididymal sperm exposed to different potentiators of DNA damage. THERIOGENOLOGY.
117, 34-39.
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Janeka, J. E., Davis, B. W., Ghosh, S., Paria, N., Das, P. J., Orlando, L., ... Raudsepp, T.
(2018).
Horse Y chromosome assembly displays unique evolutionary features and putative stallion fertility genes. Nature Communications.
9(1), 2945.
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Fisher, A., Harrison, K., Love, C., Varner, D., & Rangel, J.
(2018).
Spatio-Temporal Variation in Viability of Spermatozoa of Honey Bee, Apis mellifera, Drones in Central Texas Apiaries. SOUTHWESTERN ENTOMOLOGIST.
43(2), 343-356.
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Walbornn, S. R., Love, C. C., Blanchard, T. L., Brinsko, S. P., & Varner, D. D.
(2017).
The effect of dual-hemisphere breeding on stallion fertility. THERIOGENOLOGY.
94, 8-14.
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Serafini, R., Varner, D. D., Bissett, W., Blanchard, T. L., Teague, S. R., & Love, C. C.
(2017).
The effect of flash-freezing temperature on stallion sperm DNA structure. THERIOGENOLOGY.
95, 113-117.
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Darr, C. R., Moraes, L. E., Connon, R. E., Love, C. C., Teague, S., Varner, D. D., & Meyers, S. A.
(2017).
The relationship between mitochondrial DNA copy number and stallion sperm function. THERIOGENOLOGY.
94, 94-99.
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Glass, K. G., Arnold, C. E., Varner, D. D., Chaffin, M. K., & Schumacher, J.
(2016).
Signalment, clinical features, and outcome for male horses with urethral rents following perineal urethrotomy or corpus spongiotomy: 33 cases (1989-2013). Journal of the American Veterinary Medical Association.
249(12), 1421-1427.
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Varner, D. D.
(2016).
Strategies for Processing Semen from Subfertile Stallions for Cooled Transport. Veterinary Clinics of North America: Equine Practice.
32(3), 547-560.
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Darr, C. R., Cortopassi, G. A., Datta, S., Varner, D. D., & Meyers, S. A.
(2016).
Mitochondrial oxygen consumption is a unique indicator of stallion spermatozoal health and varies with cryopreservation media. THERIOGENOLOGY.
86(5), 1382-1392.
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Varner, D. D.
(2016).
Approaches to Breeding Soundness Examination and Interpretation of Results. Journal of Equine Veterinary Science.
43, S37-S44.
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Darr, C. R., Varner, D. D., Teague, S., Cortopassi, G. A., Datta, S., & Meyers, S. A.
(2016).
Lactate and Pyruvate Are Major Sources of Energy for Stallion Sperm with Dose Effects on Mitochondrial Function, Motility, and ROS Production. Biology of Reproduction.
95(2), 34-11.
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Turner, C. E., Walbornn, S. R., Blanchard, T. L., Varner, D. D., Brinsko, S. P., LaCaze, K. A., Teague, S. R., & Love, C. C.
(2016).
The effect of two levels of hemospermia on stallion fertility. THERIOGENOLOGY.
86(6), 1399-1402.
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Voge, J., Varner, D. D., Blanchard, T. L., Meschini, M., Turner, C., Teague, S. R., Brinsko, S. P., & Love, C. C.
(2016).
The effects of urine concentration, and cushion centrifugation to remove urine, on the quality of cool-stored stallion sperm. THERIOGENOLOGY.
86(5), 1294-1298.
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Serafini, R., Romano, J. E., Varner, D. D., Di Palo, R., & Love, C. C.
(2015).
Sperm DNA assays and their relationship to sperm motility and morphology in bulls (Bos Taurus). Animal Reproduction Science.
159, 77-86.
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Varner, D. D.
(2015).
Odyssey of the spermatozoon. Asian Journal of Andrology.
17(4), 522-528.
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Varner, D. D.
(2015).
The Ins and Outs of Spermatology. Asian Journal of Andrology.
17(4), 521-521.
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Arnold, C. E., Brinsko, S. P., & Varner, D. D.
(2015).
Cervical wedge resection for treatment of pyometra secondary to transluminal cervical adhesions in six mares. Journal of the American Veterinary Medical Association.
246(12), 1354-1357.
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Ing, N. H., Brinsko, S. P., Curley, K. O., Forrest, D. W., Love, C. C., Hinrichs, K., ... Welsh, T. H.
(2015).
Dexamethasone acutely regulates endocrine parameters in stallions and subsequently affects gene expression in testicular germ cells. Animal Reproduction Science.
152, 47-54.
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Serafini, R., Varner, D. D., Bissett, W., Blanchard, T. L., Teague, S. R., & Love, C. C.
(2015).
Sperm DNA quality evaluated by comet assay and sperm chromatin structure assay in stallions after unilateral orchiectomy. THERIOGENOLOGY.
84(5), 833-837.
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Hayden, S. S., Blanchard, T. L., Brinsko, S. P., Varner, D. D., Hinrichs, K., & Love, C. C.
(2015).
The "dilution effect" in stallion sperm. THERIOGENOLOGY.
83(4), 772-777.
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Love, C. C., Noble, J. K., Standridge, S. A., Bearden, C. T., Blanchard, T. L., Varner, D. D., & Cavinder, C. A.
(2015).
The relationship between sperm quality in cool-shipped semen and embryo recovery rate in horses. THERIOGENOLOGY.
84(9), 1587-1593.e4.
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Macas-Garca, B., Gonzlez-Fernndez, L., Loux, S. C., Rocha, A. M., Guimares, T., Pea, F. J., Varner, D. D., & Hinrichs, K.
(2015).
Effect of calcium, bicarbonate, and albumin on capacitation-related events in equine sperm. REPRODUCTION.
149(1), 87-99.
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Varner, D. D., Gibb, Z., & Aitken, R. J.
(2015).
Stallion fertility: a focus on the spermatozoon. Equine Veterinary Journal.
47(1), 16-24.
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Loux, S. C., Macas-Garcia, B., Gonzlez-Fernndez, L., Canesin, H. D., Varner, D. D., & Hinrichs, K.
(2014).
Regulation of axonemal motility in demembranated equine sperm. Biology of Reproduction.
91(6), 152-16.
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Cavinder, C. A., Perrin, A. L., Rosenberg, J. L., & Varner, D. D.
(2014).
Relative alterations in core body temperature and internal and external scrotal temperatures of exercising stallions. Professional Animal Scientist.
30(4), 451-456.
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Ing, N. H., Forrest, D. W., Love, C. C., & Varner, D. D.
(2014).
Dense spermatozoa in stallion ejaculates contain lower concentrations of mRNAs encoding the sperm specific calcium channel 1, ornithine decarboxylase antizyme 3, aromatase, and estrogen receptor alpha than less dense spermatozoa. THERIOGENOLOGY.
82(2), 347-353.
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Ing, N. H., Forrest, D. W., Riggs, P. K., Loux, S., Love, C. C., Brinsko, S. P., Varner, D. D., & Welsh, T. H.
(2014).
Dexamethasone acutely down-regulates genes involved in steroidogenesis in stallion testes. The Journal of Steroid Biochemistry and Molecular Biology.
143, 451-459.
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Kiser, A. M., Brinsko, S. P., Love, C. C., Varner, D. D., Sudderth, K., & Blanchard, T. L.
(2014).
Relationship of Sperm Quality to Fertility after 4 Days of Cooled Storage of Equine Semen. Journal of Equine Veterinary Science.
34(5), 602-605.
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Loux, S. C., Crawford, K. R., Ing, N. H., Gonzlez-Fernndez, L., Macas-Garca, B., Love, C. C., ... Hinrichs, K.
(2013).
CatSper and the relationship of hyperactivated motility to intracellular calcium and pH kinetics in equine sperm. Biology of Reproduction.
89(5), 123-15.
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Das, P. J., McCarthy, F., Vishnoi, M., Paria, N., Gresham, C., Li, G., ... Raudsepp, T.
(2013).
Stallion sperm transcriptome comprises functionally coherent coding and regulatory RNAs as revealed by microarray analysis and RNA-seq. PLoS ONE.
8(2), e56535-e56535.
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Ponthier, J., Teague, S. R., Franck, T. Y., de la Rebire, G., Serteyn, D. D., Brinsko, S. P., ... Deleuze, S. C.
(2013).
Effect of non-sperm cells removal with single-layer colloidal centrifugation on myeloperoxidase concentration in post-thaw equine semen. THERIOGENOLOGY.
80(9), 1082-1087.
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Sudderth, A. K., Kiser, A. M., Brinsko, S. P., Love, C. C., Varner, D. D., Burns, P. J., & Blanchard, T. L.
(2013).
Efficacy of Long-Acting Formulations of Estradiol or Progesterone Plus Estradiol on Estrous Synchronization in Broodmares. Journal of Equine Veterinary Science.
33(8), 670-672.
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Gonzlez-Fernndez, L., Macas-Garca, B., Loux, S. C., Varner, D. D., & Hinrichs, K.
(2013).
Focal adhesion kinases and calcium/calmodulin-dependent protein kinases regulate protein tyrosine phosphorylation in stallion sperm. Biology of Reproduction.
88(6), 138-12.
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McCormick, J. D., Valdez, R., Rakestraw, P. C., Varner, D. D., Blanchard, T. L., Cavinder, C. A., ... Love, C. C.
(2012).
Effect of surgical technique for unilateral orchiectomy on subsequent testicular function in Miniature Horse stallions. Equine Veterinary Journal.
44(43), 100-104.
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Raudsepp, T., McCue, M. E., Das, P. J., Dobson, L., Vishnoi, M., Fritz, K. L., ... Chowdhary, B. P.
(2012).
Genome-wide association study implicates testis-sperm specific FKBP6 as a susceptibility locus for impaired acrosome reaction in stallions. 8(12), e1003139-e1003139.
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Dean, C. J., Hobgood, A. M., Blodgett, G. P., Love, C. C., Blanchard, T. L., & Varner, D. D.
(2012).
The addition of ticarcillin-clavulanic acid to INRA 96 extender for stallion semen cooling. Equine Veterinary Journal.
44(43), 95-99.
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Gonzlez-Fernndez, L., Macas-Garca, B., Velez, I. C., Varner, D. D., & Hinrichs, K.
(2012).
Calcium-calmodulin and pH regulate protein tyrosine phosphorylation in stallion sperm. REPRODUCTION.
144(4), 411-422.
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Hayden, S. S., Blanchard, T. L., Brinsko, S. P., Varner, D. D., Hinrichs, K., & Love, C. C.
(2012).
Pregnancy rates in mares inseminated with 0.5 or 1 million sperm using hysteroscopic or transrectally guided deep-horn insemination techniques. THERIOGENOLOGY.
78(4), 914-920.
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Mawyer, J. D., Cavinder, C. A., Vogelsang, M. M., Sigler, D. H., Love, C. C., Brinsko, S. P., ... Gordon, R. K.
(2012).
Thermoregulation of the testicle in response to exercise and subsequent effects on semen characteristics of stallions. Journal of Animal Science.
90(8), 2532-2539.
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Bedford-Guaus, S. J., McPartlin, L. A., & Varner, D. D.
(2012).
Characterization of Equine Phospholipase C Zeta: A Review and Preliminary Results on Expression Defects in Subfertile Stallions. Journal of Equine Veterinary Science.
32(8), 445-450.
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Blanchard, T. L., Varner, D. D., Brinsko, S. P., & Love, C. C.
(2012).
Azoospermia in stallions: determining the cause. Compendium (Yardley, PA).
34(2), E2.
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Blanchard, T. L., Varner, D. D., Brinsko, S. P., & Love, C. C.
(2012).
Azoospermia in stallions: determining the cause. Compendium (Yardley, PA).
34(2),
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Blanchard, T. L., Brinsko, S. P., Love, C. C., Vest, D. D., Berezowski, C. B., Wendt, K. M., Stich, K., & Varner, D. D.
(2012).
Case Study of Processing and Insemination Techniques: Attempts to Improve Fertility of an Aged Stallion with Dilute Semen of Poor Quality. Journal of Equine Veterinary Science.
32(1), 5-11.
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Edmond, A. J., Brinsko, S. P., Love, C. C., Blanchard, T. L., Teague, S. R., & Varner, D. D.
(2012).
Effect of centrifugal fractionation protocols on quality and recovery rate of equine sperm. THERIOGENOLOGY.
77(5), 959-966.
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Love, C. C., Blanchard, T. L., Varner, D. D., Brinsko, S. P., Voge, J., Bliss, S., ... LaCaze, K.
(2012).
Effect of daily semen centrifugation and resuspension on the longevity of equine sperm quality following cooled storage. THERIOGENOLOGY.
77(9), 1911-1917.
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Blanchard, T. L., Thompson, J. A., Love, C. C., Brinsko, S. P., Ramsey, J., O'Meara, A., & Varner, D. D.
(2012).
Influence of day of postpartum breeding on pregnancy rate, pregnancy loss rate, and foaling rate in Thoroughbred mares. THERIOGENOLOGY.
77(7), 1290-1296.
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Blanchard, T. L., Varner, D. D., Love, C. C., Brinsko, S. P., & Morehead, J. P.
(2012).
Management Options for the Aged Breeding Stallion with Declining Testicular Function. Journal of Equine Veterinary Science.
32(8), 430-435.
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Bliss, S. B., Voge, J. L., Hayden, S. S., Teague, S. R., Brinsko, S. P., Love, C. C., Blanchard, T. L., & Varner, D. D.
(2012).
The impact of cushioned centrifugation protocols on semen quality of stallions. THERIOGENOLOGY.
77(6), 1232-1239.
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Olivieri, B. T., Love, B. C., Rezabek, G. B., Lamm, C. G., Varner, D. D., Payton, M. E., & Holyoak, G. R.
(2011).
Effect of Antibiotic-containing Extenders on Taylorella equigenitalis Contaminated Semen. Journal of Equine Veterinary Science.
31(11), 655-660.
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Choi, Y. H., Varner, D. D., Love, C. C., Hartman, D. L., & Hinrichs, K.
(2011).
Production of live foals via intracytoplasmic injection of lyophilized sperm and sperm extract in the horse. REPRODUCTION.
142(4), 529-538.
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Salazar, J. L., Teague, S. R., Love, C. C., Brinsko, S. P., Blanchard, T. L., & Varner, D. D.
(2011).
Effect of cryopreservation protocol on postthaw characteristics of stallion sperm. 76(3), 409-418.
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Foster, M. L., Varner, D. D., Hinrichs, K., Teague, S., Lacaze, K., Blanchard, T. L., & Love, C. C.
(2011).
Agreement between measures of total motility and membrane integrity in stallion sperm. THERIOGENOLOGY.
75(8), 1499-1505.
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Foster, M. L., Love, C. C., Varner, D. D., Brinsko, S. P., Hinrichs, K., Teague, S., Lacaze, K., & Blanchard, T. L.
(2011).
Comparison of methods for assessing integrity of equine sperm membranes. THERIOGENOLOGY.
76(2), 334-341.
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Arnold, C. E., Brinsko, S. P., Love, C. C., & Varner, D. D.
(2010).
Use of a modified Vinsot technique for partial phallectomy in 11 standing horses. Journal of the American Veterinary Medical Association.
237(1), 82-86.
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Laughlin, A. M., Welsh, T. H., Love, C. C., Varner, D. D., Parrish, A. R., Forrest, D. W., & Ing, N. H.
(2010).
In vitro culture of precision-cut testicular tissue as a novel tool for the study of responses to LH. In Vitro Cellular and Developmental Biology - Animal.
46(1), 45-53.
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Blanchard, T. L., Thompson, J. A., Brinsko, S. P., Varner, D. D., Love, C. C., Ramsey, J., & O'Meara, A.
(2010).
Pregnancy Rates in Tranquilized Maiden Thoroughbred Mares. Journal of Equine Veterinary Science.
30(4), 187-190.
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Blanchard, T. L., Thompson, J. A., Brinsko, S. P., Varner, D. D., Love, C. C., Ramsey, J., & O'Meara, A.
(2010).
Some Factors Associated With Fertility of Thoroughbred Stallions. Journal of Equine Veterinary Science.
30(8), 407-418.
-
Das, P. J., Paria, N., Gustafson-Seabury, A., Vishnoi, M., Chaki, S. P., Love, C. C., ... Raudsepp, T.
(2010).
Total RNA isolation from stallion sperm and testis biopsies. THERIOGENOLOGY.
74(6), 1099-1106e2.
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Varner, D. D.
(2008).
Developments in stallion semen evaluation. THERIOGENOLOGY.
70(3), 448-462.
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Johnson, L., Thompson, D. L., & Varner, D. D.
(2008).
Role of Sertoli cell number and function on regulation of spermatogenesis. Animal Reproduction Science.
105(1-2), 23-51.
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Waite, J. A., Love, C. C., Brinsko, S. P., Teague, S. R., Salazar, J. L., Mancill, S. S., & Varner, D. D.
(2008).
Factors impacting equine sperm recovery rate and quality following cushioned centrifugation. THERIOGENOLOGY.
70(4), 704-714.
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Varner, D. D., Love, C. C., Brinsko, S. P., Blanchard, T. L., Hartman, D. L., Bliss, S. B., Carroll, B. S., & Eslick, M. C.
(2008).
Semen Processing for the Subfertile Stallion. Journal of Equine Veterinary Science.
28(11), 677-685.
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Comerford, K. L., Love, C. C., Brinsko, S. P., Edmund, A. J., Waite, J. A., Teague, S. R., & Varner, D. D.
(2008).
Validation of a commercially available fluorescence-based instrument to evaluate stallion spermatozoal concentration. Animal Reproduction Science.
107(3-4), 316-317.
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Hinrichs, K., Choi, Y. H., Varner, D. D., & Hartman, D. L.
(2007).
Production of cloned horse foals using roscovitine-treated donor cells and activation with sperm extract and/or ionomycin. REPRODUCTION.
134(2), 319-325.
-
Brinsko, S. P., Love, C. C., Bauer, J. E., Macpherson, M. L., & Varner, D. D.
(2007).
Cholesterol-to-phospholipid ratio in whole sperm and seminal plasma from fertile stallions and stallions with unexplained subfertility. Animal Reproduction Science.
99(1-2), 65-71.
-
Choi, Y. H., Love, L. B., Varner, D. D., & Hinrichs, K.
(2007).
Effect of holding technique and culture drop size in individual or group culture on blastocyst development after ICSI of equine oocytes with low meiotic competence. Animal Reproduction Science.
102(1-2), 38-47.
-
Roasa, L. M., Choi, Y. H., Love, C. C., Romo, S., Varner, D. D., & Hinrichs, K.
(2007).
Ejaculate and type of freezing extender affect rates of fertilization of horse oocytes in vitro. THERIOGENOLOGY.
68(4), 560-566.
-
Hinrichs, K., Choi, Y. H., Walckenaer, B. E., Varner, D. D., & Hartman, D. L.
(2007).
In vitro-produced equine embryos: production of foals after transfer, assessment by differential staining and effect of medium calcium concentrations during culture. THERIOGENOLOGY.
68(4), 521-529.
-
Donnelly, C. L., Staub, C., Varner, D., Blanchard, T., Johnson, L., & Forrest, D. W.
(2007).
The effects of growth factor on testicular germ cell apoptosis in the stallion. Journal of Equine Veterinary Science.
27(5), 212-216.
-
Choi, S., Yoo, C., & Chang, O.
(2006).
Development of Integrated DAO Pattern Applying Iterator Pattern. Lecture Notes in Computer Science.
3983(4), 955-965.
-
Hinrichs, K., Choi, Y. H., Love, C. C., Chung, Y. G., & Varner, D. D.
(2006).
Production of horse foals via direct injection of roscovitine-treated donor cells and activation by injection of sperm extract. REPRODUCTION.
131(6), 1063-1072.
-
Choi, Y. H., Love, L. B., Varner, D. D., & Hinrichs, K.
(2006).
Blastocyst development in equine oocytes with low meiotic competence after suppression of meiosis with roscovitine prior to in vitro maturation. Zygote: the biology of gametes and early embryos.
14(1), 1-8.
-
Choi, Y. H., Love, C. C., Varner, D. D., & Hinrichs, K.
(2006).
Equine blastocyst development after intracytoplasmic injection of sperm subjected to two freeze-thaw cycles. THERIOGENOLOGY.
65(4), 808-819.
-
Choi, Y. H., Love, L. B., Varner, D. D., & Hinrichs, K.
(2006).
Holding immature equine oocytes in the absence of meiotic inhibitors: effect on germinal vesicle chromatin and blastocyst development after intracytoplasmic sperm injection. THERIOGENOLOGY.
66(4), 955-963.
-
Brinsko, S. P., Love, C. C., Bauer, J. E., Macpherson, M. L., & Varner, D. D.
(2005).
Cholesterol to phospholipid ratio in sperm of stallions with unexplained subfertility. Animal Reproduction Science.
89(1-4), 217-219.
-
Bosard, T., Love, C., Brinsko, S., Blanchard, T., Thompson, J., & Varner, D.
(2005).
Evaluation and diagnosis of acrosome function/dysfunction in the stallion. Animal Reproduction Science.
89(1-4), 215-217.
-
Hinrichs, K., Choi, Y. H., Love, L. B., Varner, D. D., Love, C. C., & Walckenaer, B. E.
(2005).
Chromatin configuration within the germinal vesicle of horse oocytes: changes post mortem and relationship to meiotic and developmental competence. Biology of Reproduction.
72(5), 1142-1150.
-
Pauwels, F., Schumacher, J., & Varner, D.
(2005).
Priapism in horses. Compendium (Yardley, PA).
27(4), 311-315.
-
Brinsko, S. P., Varner, D. D., Love, C. C., Blanchard, T. L., Day, B. C., & Wilson, M. E.
(2005).
Effect of feeding a DHA-enriched nutriceutical on the quality of fresh, cooled and frozen stallion semen. THERIOGENOLOGY.
63(5), 1519-1527.
-
Love, C. C., Brinsko, S. P., Rigby, S. L., Thompson, J. A., Blanchard, T. L., & Varner, D. D.
(2005).
Relationship of seminal plasma level and extender type to sperm motility and DNA integrity. THERIOGENOLOGY.
63(6), 1584-1591.
-
Lindsey, A. C., Varner, D. D., Seidel, G. E., Bruemmer, J. E., & Squires, E. L.
(2005).
Hysteroscopic or rectally guided, deep-uterine insemination of mares with spermatozoa stored 18 h at either 5 degrees C or 15 degrees C prior to flow-cytometric sorting. Animal Reproduction Science.
85(1-2), 125-130.
-
Ing, N. H., Laughlin, A. M., Varner, D. D., Welsh, T. H., Forrest, D. W., Blanchard, T. L., & Johnson, L.
(2004).
Gene expression in the spermatogenically inactive "dark" and maturing "light" testicular tissues of the prepubertal colt. Journal of Andrology.
25(4), 535-544.
-
Choi, Y. H., Roasa, L. M., Love, C. C., Varner, D. D., Brinsko, S. P., & Hinrichs, K.
(2004).
Blastocyst formation rates in vivo and in vitro of in vitro-matured equine oocytes fertilized by intracytoplasmic sperm injection. Biology of Reproduction.
70(5), 1231-1238.
-
Choi, Y. H., Love, L. B., Varner, D. D., & Hinrichs, K.
(2004).
Factors affecting developmental competence of equine oocytes after intracytoplasmic sperm injection. REPRODUCTION.
127(2), 187-194.
-
Thompson, J. A., Love, C. C., Stich, K. L., Brinsko, S. P., Blanchard, T. L., & Varner, D. D.
(2004).
A Bayesian approach to prediction of stallion daily sperm output. THERIOGENOLOGY.
62(9), 1607-1617.
-
Heninger, N. L., Staub, C., Blanchard, T. L., Johnson, L., Varner, D. D., & Forrest, D. W.
(2004).
Germ cell apoptosis in the testes of normal stallions. THERIOGENOLOGY.
62(1-2), 283-297.
-
Love, L. B., Choi, Y. H., Love, C. C., Varner, D. D., & Hinrichs, K.
(2003).
Effect of ovary storage and oocyte transport method on maturation rate of horse oocytes. THERIOGENOLOGY.
59(3-4), 765-774.
-
Brinsko, S. P., Blanchard, T. L., Rigby, S. L., Love, C. C., & Varner, D. D.
(2003).
Effects of dead spermatozoa on motion characteristics and membrane integrity of live spermatozoa in fresh and cooled-stored equine semen. 59(3-4), 735-742.
-
Choi, Y., Love, C. C., Varner, D. D., Love, L. B., & Hinrichs, K.
(2003).
Effects of gas conditions, time of medium change, and ratio of medium to embryo on in vitro development of horse oocytes fertilized by intracytoplasmic sperm injection. THERIOGENOLOGY.
59(5-6), 1219-1229.
-
Brinsko, S. P., Rigby, S. L., Lindsey, A. C., Blanchard, T. L., Love, C. C., & Varner, D. D.
(2003).
Pregnancy rates in mares following hysteroscopic or transrectally-guided insemination with low sperm numbers at the utero-tubal papilla. THERIOGENOLOGY.
59(3-4), 1001-1009.
-
Love, C. C., Thompson, J. A., Brinsko, S. P., Rigby, S. L., Blanchard, T. L., Lowry, V. K., & Varner, D. D.
(2003).
Relationship between stallion sperm motility and viability as detected by two fluorescence staining techniques using flow cytometry. THERIOGENOLOGY.
60(6), 1127-1138.
-
Lindsey, A. C., Varner, D. D., Seidel, G. E., Bruemmer, J. E., & Squires, E. L.
(2002).
Hysteroscopic or rectally guided, deep-uterine insemination of mares with spermatozoa stored 18 h at either 5 or 15 degrees C prior to flow-cytometric sorting. THERIOGENOLOGY.
58(2-4), 659-662.
-
Macpherson, M. L., Simmen, R., Simmen, F. A., Hernandez, J., Sheerin, B. R., Varner, D. D., ... Blanchard, T. L.
(2002).
Insulin-like growth factor-I and insulin-like growth factor binding protein-2 and -5 in equine seminal plasma: association with sperm characteristics and fertility. Biology of Reproduction.
67(2), 648-654.
-
Scott, M. A., Varner, D. D., Liu, I., & Enders, A. C.
(2002).
Presumptive evidence of a preovulatory sperm reservoir in the mare: morphological investigations using scanning electron microscopy. THERIOGENOLOGY.
58(2-4), 639-642.
-
Choi, Y. H., Love, C. C., Chung, Y. G., Varner, D. D., Westhusin, M. E., Burghardt, R. C., & Hinrichs, K.
(2002).
Production of nuclear transfer horse embryos by Piezo-driven injection of somatic cell nuclei and activation with stallion sperm cytosolic extract. Biology of Reproduction.
67(2), 561-567.
-
Hinrichs, K., Love, C. C., Brinsko, S. P., Choi, Y. H., & Varner, D. D.
(2002).
In vitro fertilization of in vitro-matured equine oocytes: effect of maturation medium, duration of maturation, and sperm calcium ionophore treatment, and comparison with rates of fertilization in vivo after oviductal transfer. Biology of Reproduction.
67(1), 256-262.
-
Choi, Y. H., Love, C. C., Love, L. B., Varner, D. D., Brinsko, S., & Hinrichs, K.
(2002).
Developmental competence in vivo and in vitro of in vitro-matured equine oocytes fertilized by intracytoplasmic sperm injection with fresh or frozen-thawed spermatozoa. 123(3), 455-465.
-
Hinrichs, K., Love, C. C., Choi, Y. H., Varner, D. D., Wiggins, C. N., & Reinoehl, C.
(2002).
Suppression of meiosis by inhibitors of m-phase proteins in horse oocytes with low meiotic competence. Zygote: the biology of gametes and early embryos.
10(1), 37-45.
-
Choi, Y., Shin, T., Love, C. C., Johnson, C., Varner, D. D., Westhusin, M. E., & Hinrichs, K.
(2002).
Effect of co-culture with theca interna on nuclear maturation of horse oocytes with low meiotic competence, and subsequent fusion and activation rates after nuclear transfer. THERIOGENOLOGY.
57(3), 1005-1011.
-
Love, C. C., Love, L. B., Varner, D. D., & Hinrichs, K.
(2002).
Effect of holding at room temperature on initial chromatin configuration and in vitro maturation rate of equine oocytes. THERIOGENOLOGY.
57(8), 1973-1979.
-
Lo, C. C., Thompson, J. A., Lowry, V. K., & Varner, D. D.
(2002).
Effect of storage time and temperature on stallion sperm DNA and fertility. THERIOGENOLOGY.
57(3), 1135-1142.
-
Blanchard, T. L., Woods, J. A., Brinsko, S. P., Varner, D. D., & Boothe, D. M.
(2002).
Theriogenology question of the month. Journal of the American Veterinary Medical Association.
221(6), 793-795.
-
Choi, Y. H., Love, C. C., Chung, Y. G., Varner, D. D., Westhusin, M. E., Burghardt, R. C., & Hinrichs, K.
(2002).
Use of Piezo-driven direct nuclear injection and activation with stallion sperm extract to produce horse nuclear transfer embryos. THERIOGENOLOGY.
58(2-4), 771-774.
-
Macpherson, M. L., Blanchard, T. L., Love, C. C., Brinsko, S. P., & Varner, D. D.
(2002).
Use of a silane-coated silica particle solution to enhance the quality of ejaculated semen in stallions. THERIOGENOLOGY.
58(2-4), 317-320.
-
Rigby, S. L., Barhoumi, R., Burghardt, R. C., Colleran, P., Thompson, J. A., Varner, D. D., ... Delp, M. D.
(2001).
Mares with delayed uterine clearance have an intrinsic defect in myometrial function. Biology of Reproduction.
65(3), 740-747.
-
Rammerstorfer, C., Potter, G. D., Brumbaugh, G. W., Gibbs, P. G., Varner, D. D., & Rammerstorfer, E. H.
(2001).
Physiologic responses of acclimatized or non-acclimatized mature reining horses to heat stress: I. Heart rate, respiration rate, lactate, rectal temperature, cortisol and packed cell volume. Journal of Equine Veterinary Science.
21(9), 431-438.
-
Choi, Y. H., Love, C. C., Varner, D. D., Thompson, J. A., & Hinrichs, K.
(2001).
Activation of cumulus-free equine oocytes: effect of maturation medium, calcium ionophore concentration and duration of cycloheximide exposure. REPRODUCTION.
122(1), 177-183.
-
Rigby, S. L., Brinsko, S. P., Cochran, M., Blanchard, T. L., Love, C. C., & Varner, D. D.
(2001).
Advances in cooled semen technologies: seminal plasma and semen extender. Animal Reproduction Science.
68(3-4), 171-180.
-
Blanchard, T. L., Johnson, L., Varner, D. D., Rigby, S. L., Brinsko, S. P., Love, C. C., & Miller, C.
(2001).
Low daily sperm output per ml of testis as a diagnostic criteria for testicular degeneration in stallions. Journal of Equine Veterinary Science.
21(1), 11-+.
-
Blanchard, T., Varner, D., Miller, C., & Roser, J.
(2000).
Recommendations for clinical GnRH challenge testing of stallions. Journal of Equine Veterinary Science.
20(11), 678-737.
-
Johnson, L., Varner, D. D., Roberts, M. E., Smith, T. L., Keillor, G. E., & Scrutchfield, W. L.
(2000).
Efficiency of spermatogenesis: a comparative approach. Animal Reproduction Science.
60-61, 471-480.
-
Brinsko, S. P., Rowan, K. R., Varner, D. D., & Blanchard, T. L.
(2000).
Effects of transport container and ambient storage temperature on motion characteristics of equine spermatozoa. THERIOGENOLOGY.
53(8), 1641-1655.
-
Woods, J., Rigby, S., Brinsko, S., Stephens, R., Varner, D., & Blanchard, T.
(2000).
Effect of intrauterine treatment with prostaglandin E2 prior to insemination of mares in the uterine horn or body. THERIOGENOLOGY.
53(9), 1827-1836.
-
Varner, D. D., Blanchard, T. L., Brinsko, S. P., Love, C. C., Taylor, T. S., & Johnson, L.
(2000).
Techniques for evaluating selected reproductive disorders of stallions. Animal Reproduction Science.
60-61, 493-509.
-
Combes, G. B., Varner, D. D., Schroeder, F., Burghardt, R. C., & Blanchard, T. L.
(2000).
Effect of cholesterol on the motility and plasma membrane integrity of frozen equine spermatozoa after thawing. Journal of reproduction and fertility. Supplement.
127-132.
-
Bedford, S. J., Varner, D. D., & Meyers, S. A.
(2000).
Effects of cryopreservation on the acrosomal status of stallion spermatozoa. Journal of reproduction and fertility. Supplement.
133-140.
-
Love, C. C., Varner, D. D., & Thompson, J. A.
(2000).
Intra- and inter-stallion variation in sperm morphology and their relationship with fertility. Journal of reproduction and fertility. Supplement.
93-100.
-
Evenson, D. P., Jost, L. K., & Varner, D. D.
(2000).
Relationship between sperm nuclear protamine free -SH status and susceptibility to DNA denaturation. Journal of reproduction and fertility. Supplement.
401-406.
-
Blanchard, T., Varner, D., Johnson, L., Roser, J., Hill, J., & Miller, C.
(2000).
Testicular and hormonal changes in stallions with thermally induced testicular degeneration. Journal of reproduction and fertility. Supplement.
51-59.
-
Rigby, S., Hill, J., Miller, C., Thompson, J., Varner, D., & Blanchard, T.
(1999).
Administration of oxytocin immediately after insemination does not improve pregnancy rates in mares bred by fertile or subfertile stallions. THERIOGENOLOGY.
51(6), 1143-1150.
-
Schumacher, J., Varner, D. D., Crabill, M. R., & Blanchard, T. L.
(1999).
The effect of a surgically created shunt between the corpus cavernosum penis and corpus spongiosum penis of stallions on erectile and ejaculatory function. Veterinary Surgery.
28(1), 21-24.
-
Hill, J., Westhusin, M., & Varner, D. D.
(1998).
Advances in large-animal reproduction: In vitro production of embryos. VETERINARY MEDICINE.
93(6), 575-+.
-
Hill, J., Westhusin, M., & Varner, D. D.
(1998).
Advances in large-animal reproduction: The expanding role of ultrasonography. VETERINARY MEDICINE.
93(5), 480-484.
-
Hill, J., Westhusin, M., & Varner, D. D.
(1998).
Advances in large-animal reproduction: Updated techniques for artificial insemination and embryo transfer. VETERINARY MEDICINE.
93(5), 485-490.
-
Blanchard, T. L., & Varner, D. D.
(1998).
Breeding with cooled, transported equine semen: Proper mare management. VETERINARY MEDICINE.
93(7), 665-671.
-
Blanchard, T. L., & Varner, D. D.
(1998).
Breeding with cooled, transported equine semen: Testing and preparing semen for travel. VETERINARY MEDICINE.
93(5), 474-479.
-
Blanchard, T. L., & Varner, D. D.
(1998).
Breeding with cooled, transported equine semen: The insemination process. VETERINARY MEDICINE.
93(9), 831-837.
-
Shore, M. D., Macpherson, M. L., Combes, G. B., Varner, D. D., & Blanchard, T. L.
(1998).
Fertility comparison between breeding at 24 hours or at 24 and 48 hours after collection with cooled equine semen. THERIOGENOLOGY.
50(5), 693-698.
-
Varner, D. D., Scanlan, C. M., Thompson, J. A., Brumbaugh, G. W., Blanchard, T. L., Carlton, C. M., & Johnson, L.
(1998).
Bacteriology of preserved stallion semen and antibiotics in semen extenders. THERIOGENOLOGY.
50(4), 559-573.
-
Blanchard, T. L., & Varner, D. D.
(1998).
Breeding with cooled transported equine semen: The advantages and success rate. VETERINARY MEDICINE.
93(3), 275-279.
-
Rammerstorfer, C., Potter, G. D., Cudd, T. A., Gibbs, P. G., Varner, D. D., & Householder, D. D.
(1998).
Physiological responses of mature quarter horses to reining training when fed conventional and fat-supplemented diets. 18(3), 175-183.
-
Rigby, S., Love, C., Carpenter, K., Varner, D., & Blanchard, T.
(1998).
Use of prostaglandin E2 to ripen the cervix of the mare prior to induction of parturition. THERIOGENOLOGY.
50(6), 897-904.
-
Blanchard, T. L., & Varner, D. D.
(1997).
Evaluating breeding soundness in stallions .5. Predicting potential fertility. VETERINARY MEDICINE.
92(9), 815-818.
-
Katila, T., Combes, G. B., Varner, D. D., & Blanchard, T. L.
(1997).
Comparison of three containers used for the transport of cooled stallion semen. THERIOGENOLOGY.
48(7), 1085-1092.
-
Johnson, L., Blanchard, T. L., Varner, D. D., & Scrutchfield, W. L.
(1997).
Factors affecting spermatogenesis in the stallion. THERIOGENOLOGY.
48(7), 1199-1216.
-
Macpherson, M. L., Chaffin, M. K., Carroll, G. L., Jorgensen, J., Arrott, C., Varner, D. D., & Blanchard, T. L.
(1997).
Three methods of oxytocin-induced parturition and their effects of foals. Journal of the American Veterinary Medical Association.
210(6), 799-803.
-
Blanchard, T. L., & Varner, D. D.
(1996).
Evaluating breeding soundness in stallions .3. Useful laboratory tests. VETERINARY MEDICINE.
91(3), 263-267.
-
Blanchard, T. L., Varner, D. D., & Brinsko, S. P.
(1996).
Theriogenology question of the month. Scrotal hematocele. Journal of the American Veterinary Medical Association.
209(12), 2013-2014.
-
Blanchard, T. L., Jorgensen, J. B., Varner, D. D., Forrest, D. W., & Evans, J. W.
(1996).
Clinical observations on changes in concentrations of hormones in plasma of two stallions with thermally-induced testicular degeneration. Journal of Equine Veterinary Science.
16(5), 195-201.
-
Blanchard, T. L., & Varner, D. D.
(1996).
Evaluating breeding soundness in stallions .4. Hormonal assay and testicular biopsy. VETERINARY MEDICINE.
91(4), 358-+.
-
Blanchard, T. L., & Varner, D. D.
(1996).
Evaluating breeding soundness in stallions .1. The basic examination. VETERINARY MEDICINE.
91(1), 54-&.
-
Blanchard, T. L., Varner, D. D., & Brinsko, S. P.
(1996).
Theriogenology question of the month. Journal of the American Veterinary Medical Association.
209(12), 2013-2014.
-
Schumacher, J., Varner, D. D., Schmitz, D. G., & Blanchard, T. L.
(1995).
Urethral defects in geldings with hematuria and stallions with hemospermia. Veterinary Surgery.
24(3), 250-254.
-
BLANCHARD, T. L., & VARNER, D. D.
(1995).
MANIPULATING ESTRUS IN THE MARE .1. VETERINARY MEDICINE.
90(2), 180-184.
-
BLANCHARD, T. L., & VARNER, D. D.
(1995).
MANIPULATING ESTRUS IN THE MARE .2. VETERINARY MEDICINE.
90(3), 285-289.
-
BLANCHARD, T. L., MARTIN, M. T., VARNER, D. D., & MACPHERSON, M. L.
(1995).
PREPARING FOR PARTURITION IN THE MARE. VETERINARY MEDICINE.
90(6), 597-&.
-
Arrott, C., Macpherson, M., Blanchard, T., Varner, D., Thompson, J., Simpson, B., ... Burns, P.
(1994).
Biodegradable estradiol microspheres do not affect uterine involution or characteristics of postpartum estrus in mares. THERIOGENOLOGY.
42(2), 371-384.
-
MCCARTHY, P. F., HOOPER, R. N., CARTER, G. K., VARNER, D. D., TAYLOR, T. S., & BLANCHARD, T. L.
(1994).
POSTPARTURIENT HEMORRHAGE IN THE MARE - MANAGING RUPTURED ARTERIES OF THE BROAD LIGAMENT. VETERINARY MEDICINE.
89(2), 147-152.
-
Blanchard, T. L., Varner, D. D., Fernandez, M., Buoen, L., Zhang, T. Q., Womack, J. E., & Owens, E.
(1994).
Stallion karyotype variability and lack of association with abortion: a case report. THERIOGENOLOGY.
41(4), 777-784.
-
Johnson, L., Carter, G. K., Varner, D. D., Taylor, T. S., Blanchard, T. L., & Rembert, M. S.
(1994).
The relationship of daily sperm production with number of Sertoli cells and testicular size in adult horses: role of primitive spermatogonia. Reproduction.
100(1), 315-321.
-
Varner, D. D., Bowen, J. A., & Johnson, L.
(1993).
Effect of heparin on capacitation/acrosome reaction of equine sperm. Systems Biology in Reproductive Medicine.
31(3), 199-207.
-
Hooper, R. N., Taylor, T. S., Varner, D. D., & Blanchard, T. L.
(1993).
Effects of bilateral ovariectomy via colpotomy in mares: 23 cases (1984-1990). Journal of the American Veterinary Medical Association.
203(7), 1043-1046.
-
ELLINGTON, J. E., VARNER, D. D., BURGHARDT, R. C., MEYERSWALLEN, V. N., BARHOUMI, R., BRINSKO, S. P., & BALL, B. A.
(1993).
CELL-TO-CELL COMMUNICATION OF EQUINE UTERINE TUBE (OVIDUCT) CELLS AS DETERMINED BY ANCHORED CELL ANALYSIS IN CULTURE. Animal Reproduction Science.
30(4), 313-324.
-
VARNER, D. D., BOWEN, J. A., & JOHNSON, L.
(1993).
EFFECT OF HEPARIN ON CAPACITATION/ACROSOME REACTION OF EQUINE SPERM. Systems Biology in Reproductive Medicine.
31(3), 199-207.
-
Ellington, J. E., Ignotz, G. G., Varner, D. D., Marcucio, R. S., Mathison, P., & Ball, B. A.
(1993).
In vitro interaction between oviduct epithelial and equine sperm. Systems Biology in Reproductive Medicine.
31(2), 79-86.
-
Hooper, R. N., Taylor, T. S., Behrens, E. A., & Varner, D. D.
(1992).
Use of an ecraseur for ovariohysterectomy in mares. Veterinary Surgery.
21(5), 374-377.
-
Brinsko, S. P., Varner, D. D., Blanchard, T. L., Relford, R. L., & Johnson, L.
(1992).
Bilateral infectious epididymitis in a stallion. Equine Veterinary Journal.
24(4), 325-328.
-
Brinsko, S. P., & Varner, D. D.
(1992).
Artificial insemination and preservation of semen. Veterinary Clinics of North America: Equine Practice.
8(1), 205-218.
-
Blanchard, T. L., Varner, D. D., Burns, P. J., Everett, K. A., Brinsko, S. P., & Boehnke, L.
(1992).
Regulation of estrus and ovulation in mares with progesterone or progesterone and estradiol biodegradable microspheres with or without PGF2alpha. THERIOGENOLOGY.
38(6), 1091-1106.
-
Varner, D. D., Bowen, J. A., & Johnson, L.
(1992).
Capacitation and acrosome reaction of equine spermatozoa by heparin. 4(1-2), 81-99.
-
Ellington, J. E., Varner, D. D., Mathison, P., Ignotz, G. G., Padilla, A. W., Miller, P. G., Marcucio, R. S., & Ball, B. A.
(1992).
Equine sperm and oviductal epithelia (uterine tubal) interactions in co- culture. 4(1-2), 101-111.
-
Bretzlaff, K. N., Nuti, L. C., Scarfe, A. D., Elmore, R. G., Capehart, J., Varner, D. D., & Weston, P. G.
(1991).
Luteinizing hormone and progesterone concentrations and induction of estrus after use of norgestomet ear implants or constant infusion of gonadotropin-releasing hormone in anestrous, nonlactating dairy goats. American Journal of Veterinary Research.
52(9), 1423-1426.
-
Johnson, L., Varner, D. D., Tatum, M. E., & Scrutchfield, W. L.
(1991).
Season but not age affects Sertoli cell number in adult stallions. Biology of Reproduction.
45(3), 404-410.
-
BRINSKO, S. P., VARNER, D. D., & BLANCHARD, T. L.
(1991).
THE EFFECT OF UTERINE LAVAGE PERFORMED 4 HOURS POST INSEMINATION ON PREGNANCY RATE IN MARES. THERIOGENOLOGY.
35(6), 1111-1119.
-
Meyers, P. J., & Varner, D. D.
(1991).
Abortion of a mummified fetus associated with short uterine body in a mare. Journal of the American Veterinary Medical Association.
198(10), 1768-1770.
-
Blanchard, T. L., Schumacher, J., Edwards, J. F., Varner, D. D., Lewis, R. D., Everett, K., & Joyce, J. R.
(1991).
Priapism in a stallion with generalized malignant melanoma. Journal of the American Veterinary Medical Association.
198(6), 1043-1044.
-
Varner, D. D., Vaughan, S. D., & Johnson, L.
(1991).
Use of a computerized system for evaluation of equine spermatozoal motility. American Journal of Veterinary Research.
52(2), 224-230.
-
Blanchard, T. L., Varner, D. D., Brinsko, S. P., Quirk, K., Rugila, J. N., & Boehnke, L.
(1991).
Effects of ecbolic agents on measurements of uterine involution in the mare. THERIOGENOLOGY.
36(4), 559-571.
-
Varner, D. D., Forrest, D. W., Fuentes, F., Taylor, T. S., Hooper, R. N., Brinsko, S. P., & Blanchard, T. L.
(1991).
Measurements of glycosaminoglycans in follicular, oviductal and uterine fluids of mares. Journal of reproduction and fertility. Supplement.
44, 297-306.
-
Watkins, J. P., Taylor, T. S., Day, W. C., Varner, D. D., Schumacher, J., Baird, A. N., & Welch, R. D.
(1990).
Elective cesarean section in mares: eight cases (1980-1989). Journal of the American Veterinary Medical Association.
197(12), 1639-1645.
-
VARNER, D. D., & BLANCHARD, T. L.
(1990).
AN UPDATE ON UTERINE DEFENSE-MECHANISMS IN THE MARE. Journal of Equine Veterinary Science.
10(3), 169-175.
-
Brinsko, S. P., Varner, D. D., Blanchard, T. L., & Meyers, S. A.
(1990).
The effect of postbreeding uterine lavage on pregnancy rate in mares. THERIOGENOLOGY.
33(2), 465-475.
-
BLANCHARD, T. L., EVANS, J. W., VARNER, D. D., MOLLETT, T. A., HARDIN, D. K., ELMORE, R. G., & YOUNGQUIST, R. S.
(1990).
PULSATILE RELEASE OF GONADOTROPINS IN YOUNG PONY STALLIONS. THERIOGENOLOGY.
34(6), 1087-1097.
-
Varner, D. D., Blanchard, T. L., Meyers, P. J., & Meyers, S. A.
(1989).
Fertilizing capacity of equine spermatozoa stored for 24 hours at 5 or 20 degrees C. THERIOGENOLOGY.
32(4), 515-525.
-
Althouse, G. C., Seager, S. W., Varner, D. D., & Webb, G. W.
(1989).
Diagnostic aids for the detection of urine in the equine ejaculate. THERIOGENOLOGY.
31(6), 1141-1148.
-
Blanchard, T. L., Varner, D. D., Brinsko, S. P., Meyers, S. A., & Johnson, L.
(1989).
Effects of postparturient uterine lavage on uterine involution in the mare. THERIOGENOLOGY.
32(4), 527-535.
-
Johnson, L., & Varner, D. D.
(1988).
Effect of daily spermatozoan production but not age on transit time of spermatozoa through the human epididymis. Biology of Reproduction.
39(4), 812-817.
-
Memon, M. A., Usenik, E. A., Varner, D. D., & Meyers, P. J.
(1988).
Penile paralysis and paraphimosis associated with reserpine administration in a stallion. THERIOGENOLOGY.
30(2), 411-419.
-
MEYERS, P. J., ELMORE, R. G., VARNER, D. D., BLANCHARD, T. L., SHULL, J. W., & TODD, J.
(1988).
USE OF A RAPID PROGESTERONE ASSAY IN A BEEF-CATTLE ESTRUS SYNCHRONIZATION PROGRAM. THERIOGENOLOGY.
29(6), 1285-1294.
-
Blanchard, T. L., Varner, D. D., Hurtgen, J. P., Love, C. C., Cummings, M. R., Strezmienski, P. J., Benson, C., & Kenney, R. M.
(1988).
Bilateral seminal vesiculitis and ampullitis in a stallion. Journal of the American Veterinary Medical Association.
192(4), 525-526.
-
Kloppe, L. H., Varner, D. D., Elmore, R. G., Bretzlaff, K. N., & Shull, J. W.
(1988).
Effect of insemination timing on the fertilizing capacity of frozen/thawed equine spermatozoa. THERIOGENOLOGY.
29(2), 429-439.
-
Varner, D. D., Blanchard, T. L., Love, C. L., Garcia, M. C., & Kenney, R. M.
(1988).
Effects of cooling rate and storage temperature on equine spermatozoal motility parameters. THERIOGENOLOGY.
29(5), 1043-1054.
-
Varner, D. D., Blanchard, T. L., Love, C. L., Garcia, M. C., & Kenney, R. M.
(1987).
Effects of semen fractionation and dilution ratio on equine spermatozoal motility parameters. THERIOGENOLOGY.
28(5), 709-723.
-
Varner, D. D., Ward, C. R., Storey, B. T., & Kenney, R. M.
(1987).
Induction and characterization of acrosome reaction in equine spermatozoa. American Journal of Veterinary Research.
48(9), 1383-1389.
-
Blanchard, T. L., Varner, D. D., Love, C. C., Hurtgen, J. P., Cummings, M. R., & Kenney, R. M.
(1987).
Use of a semen extender containing antibiotic to improve the fertility of a stallion with seminal vesiculitis due to Pseudomonas aeruginosa. THERIOGENOLOGY.
28(4), 541-546.
-
Strzemienski, P. J., Sertich, P. L., Varner, D. D., & Kenney, R. M.
(1987).
Evaluation of cellulose acetate/nitrate filters for the study of stallion sperm motility. Journal of reproduction and fertility. Supplement.
35, 33-38.
-
Blanchard, T. L., Varner, D. D., Buonanno, A. M., Palmer, J. E., Hansen, T. O., & Divers, T. J.
(1987).
Hydrallantois in two mares. Journal of Equine Veterinary Science.
7(4), 222-225.
-
Van Huffel, X. M., Varner, D. D., Hinrichs, K., Garcia, M. C., Strzemienski, P. J., & Kenney, R. M.
(1985).
Photomicrographic evaluation of stallion spermatozoal motility characteristics. American Journal of Veterinary Research.
46(6), 1272-1275.
-
Varner, D. D., Hinrichs, K., Garcia, M. C., Osborne, H. G., Blanchard, T. L., & Kenney, R. M.
(1985).
A comparison between cervical dimensions of pregnant and nonpregnant Santa Gertrudis and Bos taurus cows. THERIOGENOLOGY.
24(1), 109-118.
-
Brown, J. S., Varner, D. D., Hinrichs, K., & Kenney, R. M.
(1984).
Surgical repair of the lacerated cervix in the mare. THERIOGENOLOGY.
22(4), 351-359.
-
Serafini, R., Ghosh, S., Love, C. C., Medrano, J., Teague, S. R., LaCaze, K. A., & Varner, D. D.
(2019).
Effect of artificial vagina lubricants on stallion sperm quality. THERIOGENOLOGY.
139, 121-125.
-
book
-
Brinsko, S., Blanchard, T., Varner, D., Schumacher, J., & Love, C.
(2011).
Manual of Equine Reproduction. Elsevier.
-
Brinsko, S., Blanchard, T., Varner, D., Schumacher, J., & Love, C.
(2011).
Manual of Equine Reproduction. Elsevier.
-
chapter
-
Blanchard, T. L., Varner, D. D., & Morehead, J. P.
(2015).
Breeding Management of the Older Stallion With Declining Testicular Function. Robinson's Current Therapy in Equine Medicine.
(pp. 641-643).
Elsevier.
-
Varner, D. D.
(2015).
Low Sperm Count. Robinson's Current Therapy in Equine Medicine.
(pp. 653-657).
Elsevier.
-
Schumacher, J., & Varner, D. D.
(2007).
Surgical Correction of Abnormalities Affecting the Reproductive Organs of Stallions. Current Therapy in Large Animal Theriogenology.
(pp. 23-36).
-
SCHUMACHER, J., & VARNER, D. D.
(2007).
Surgical Correction of Abnormalities Affecting the Reproductive Organs of Stallions. Current Therapy in Large Animal Theriogenology.
(pp. 23-36).
Elsevier.
-
VARNER, D.
(2003).
Liquid Preservation of Equine Semen. Current Therapy in Equine Medicine.
(pp. 266-269).
Elsevier.
-
Blanchard, T. L., Varner, D. D., & Morehead, J. P.
(2015).
Breeding Management of the Older Stallion With Declining Testicular Function. Robinson's Current Therapy in Equine Medicine.
(pp. 641-643).
Elsevier.
-
conference paper
-
Love, C. C., White, R. D., & Varner, D. D.
(2005).
Prepackaging of equine semen in goblets prior to cryopreservation. Animal Reproduction Science.
248-250.
-
Heninger, N., Staub, C., Blanchard, T., Varner, D., Love, C., Forrest, D., & Johnson, L.
(2002).
Apoptosis as a mechanism of germ cell loss in mature stallions. THERIOGENOLOGY.
373-376.
-
Staub, C., Heninger, N. L., Donnelly, C. L., Forrest, D. W., Blanchard, T. L., Love, C. C., Varner, D. D., & Johnson, L.
(2002).
Apoptosis during the initiation of equine spermatogenesis. THERIOGENOLOGY.
367-371.
-
Laughlin, A. M., Forrest, D. W., Varner, D. D., Blanchard, T. L., Love, C. C., Welsh, T. H., Johnson, L., & Ing, N. H.
(2002).
DNA microarray analysis of gene expression in testicular tissue of stallions. THERIOGENOLOGY.
413-415.
-
Wendt, K. M., Love, C. C., Brinsko, S. P., Thompson, J. A., Blanchard, T. L., & Varner, D. D.
(2002).
Effect of extender pH on motility characteristics of cool-stored equine spermatozoa. THERIOGENOLOGY.
321-324.
-
Hinrichs, K., Choi, Y. H., Love, L. B., Love, C. C., Varner, D. D., & Ingram, L. A.
(2002).
Effects of holding time and media on meiotic and developmental competence of horse oocytes. THERIOGENOLOGY.
675-678.
-
Donnelly, C., Staub, C., Forrest, D., Varner, D., Blanchard, T., Love, C., & Johnson, L.
(2002).
Granulocyte-macrophage colony-stimulating factor reduces testicular germ cell apoptosis in vitro. THERIOGENOLOGY.
377-380.
-
Varner, D. D., Thompson, J. A., Blanchard, T. L., Heck, R., Love, C. C., Brinsko, S. P., & Johnson, L.
(2002).
Induction of the acrosome reaction in stallion spermatozoa: effects of incubation temperature, incubation time, and ionophore concentration. THERIOGENOLOGY.
303-306.
-
Love, C. C., Thompson, J. A., Brinsko, S. P., Rigby, S. L., Blanchard, T. L., & Varner, D. D.
(2002).
Relationship of seminal plasma level and extender type to sperm motility and DNA integrity. THERIOGENOLOGY.
221-224.
-
Stich, K., Brinsko, S., Thompson, J., Love, C., Miller, C., Blanchard, T., & Varner, D.
(2002).
Stabilization of extragonadal sperm reserves in stallions: application for determination of daily sperm output. THERIOGENOLOGY.
397-400.
-
Blanchard, T. L., & Varner, D. D.
(1996).
Evaluating breeding soundness in stallions .2. Semen collection and evaluation. VETERINARY MEDICINE.
144-155.
-
Johnson, L., Varner, D. D., & Thompson, D. L.
(1991).
Effect of age and season on the establishment of spermatogenesis in the horse. Journal of reproduction and fertility. Supplement.
87-97.
-
Love, C. C., White, R. D., & Varner, D. D.
(2005).
Prepackaging of equine semen in goblets prior to cryopreservation. Animal Reproduction Science.
248-250.
Teaching
teaching activities
- VLCS485 Directed Studies Instructor
- VLCS685 Directed Studies Instructor
- VLCS691 Research Instructor
Works By Students
chaired theses and dissertations
-
Bosard, Tegan S. (2009-06). Flow cytometric evaluation of acrosome function/dysfunction in the stallion.
-
Edmond, Ann J. (2010-07). Effect of Density Gradient Centrifugation on Quality and Recovery Rate of Equine Sperm.
-
Heninger, Noah Leland, III (2007-04). The role of testicular germ cell apoptosis during equine spermatogenesis.
-
Salazar, Jose L. (2010-10). Effect of Cryopreservation Protocol on Post-Thaw Characteristics of Stallion Spermatozoa.
-
Stump, Karen Elizabeth (2013-04). Evaluation of Stallion Sperm Membrane Integrity Using Varied Flow Cytometer-Based Methodologies.
-
Waite, Jessica Arlene (2009-05). Cushioned centrifugation of stallion semen: factors impacting equine sperm recovery rate and quality.
-
Whigham, Alison R (2013-09). Effect of Seminal Plasma on Equine Sperm Quality: Preparation and Storage Techniques.
Background
education and training
- M.S. in Veterinary Anatomy, Texas A&M University - (College Station, Texas, United States) 1990
- Cert. in Animal Reproduction Residency, University of Pennsylvania - (Philadelphia, Pennsylvania, United States) 1983
- D.V.M. in Veterinary Medicine, University of Missouri - (Columbia, Missouri, United States) 1978
- B.S. in Agriculture, University of Missouri - (Columbia, Missouri, United States) 1976
awards and honors
Contact
first name
- Dickson
has last name
- Varner
mailing address
-
Texas A&M University
Large Animal Clinical Sciences
4475 TAMU
College Station, TX 77843-4475
USA
Other
In the News
- dvarner@tamu.edu